MOSCOW-A Russian reconnaissance aircraft was shot down by Syria forces responding to an Israeli airstrike, killing all 15 people aboard, in what President Vladimir Putin said Tuesday was "a chain of tragic accidental circumstances."
The downing of the Il-20 highlighted the dangers posed by the conflicting interests of various powers in the crowded skies over Syria and threatened the close security ties between Russia and Israel.
In an effort to maintain that relationship, Israeli Prime Minister Benjamin Netanyahu quickly called Putin to express sorrow over the death of the plane's crew, blamed the plane's loss squarely on Syria and offered to send Israel's air force chief to Moscow to share information about the incident.
The Russian military said the plane was hit 35 kilometers (22 miles) offshore late Monday night as it was returning to the Russian air base in Syria.
The incident triggered testy exchanges of blame between Israel and Russia.
The Israeli military said its fighter jets were targeting a Syrian military facility involved in providing weapons for Iran's proxy Hezbollah militia, noting that it warned Russia of the coming raid in line with de-confliction agreements. It said the Syrian army launched the missiles that hit the plane when the Israeli jets were already inside Israeli airspace.
But the Russian Defense Ministry said the Israeli warning came less than a minute before the strike, leaving the Russian aircraft in the line of fire. It pointedly accused the Israeli military of deliberately using the Russian plane as a cover to dodge the Syrian defenses and threatened to retaliate.
"The Israeli pilots were using the Russian aircraft as a shield and pushed it into the line of fire of the Syrian air defense," said Russian Defense Ministry spokesman Maj. Gen. Igor Konashenkov.
Russian Defense Minister Sergei Shoigu called his Israeli counterpart, Avigdor Lieberman, to declare that "the Israeli side bears full responsibility" for the plane's downing and to warn that Russia "reserves the right to retaliate."
But Putin took a more cautious tone, describing the incident as "a chain of tragic accidental circumstances." At the same time, he said Russia will respond by "taking additional steps to protect our servicemen and assets in Syria."
"It will be the steps that everyone will notice," he said without elaboration.
Netanyahu, who has maintained warm personal ties with Putin and frequently traveled to Russia for Syria-focused talks, noted the need for Russia and Israel to continue coordinating their action in Syria. At the same time, he emphasized Israel would not tolerate the Iranian military presence in Syria.